France head coach Guy Noves has recalled Stade Francais fly-half Jules Plisson for Les Bleus' summer Tests against the Springboks.
Plisson has failed to feature since the tour of Argentina last summer, but has returned to the squad on the back of strong performances for Stade in the Top 14 and Challenge Cup.
La Rochelle duo Mohamed Boughanmi and Vincent Rattez, as well as Damian Penaud (Clermont) and Anthony Jelonch (Castres) are all in line to make their debuts during the three-Test series.
Meanwhile, Stade fullback Djibril Camara has been replaced in the squad by another new face, Bordeaux-Begles wing Nans Ducuing.
Forwards: Uini Atonio, Eddy Ben Arous, Yacouba Camara, Camille Chat, Xavier Chiocci, Loann Goujon, Kevin Gourdon, Guilhem Guirado, Arthur Iturria, Anthony Jelonch, Julien Le Devedec, Bernard Le Roux, Clermont Maynadier, Yoann Maestri, Louis Picamoles, Rabah Slimani, Romain Taofifenua, Jefferson Poirot.
Backs: Henry Chavancy, Jonathan Danty, Jean Marc Doussain, Nans Ducuing, Brice Dulin, Antoine Dupont, Gael Fickou, Yoann Huget, Camille Lopez, Maxime Machenaud, Damian Penaud, Jules Plisson, Vincent Rattez, Baptiste Serin, Scott Spedding, Virimi Vakatawa.