Scotland will be without London Irish duo Sean Maitland and Blair Cowan for Saturday's Six Nations match in Italy.
Maitland misses out with a thigh injury while Cowan is sidelined with concussion. They are two of five changes made by Vern Cotter wiith Alex Allan, Chris Fusaro and Pat MacArthur also left out of the squad due to injury.
Glasgow Warriors trio Grayson Hart (scrum-half), Rob Harley and Ryan Wilson (back-row), join Edinburgh Rugby's uncapped hooker George Turner and back-row Hamish Watson in the squad while Scotland also have Tim Visser and Peter Horne available after spells out with injury.
Scotland squad for Italy:
Hooker: Ross Ford, Stuart McInally, George Turner (all Edinburgh Rugby)
Front-row: Alasdair Dickinson (Edinburgh Rugby), Zander Fagerson (Glasgow Warriors), Moray Low (Exeter Chiefs), Willem Nel (Edinburgh Rugby), Gordon Reid (Glasgow Warriors), Rory Sutherland (Edinburgh Rugby)
Lock: Jonny Gray (Glasgow Warriors), Richie Gray (Castres), Tim Swinson (Glasgow Warriors), Ben Toolis (Edinburgh Rugby)
Back-row: John Barclay (Scarlets), David Denton (Bath), John Hardie (Edinburgh Rugby), Rob Harley, Josh Strauss, Ryan Wilson (Glasgow Warriors), Hamish Watson (Edinburgh Rugby)
Scrum-half: Sam Hidalgo-Clyne (Edinburgh Rugby), Grayson Hart (Glasgow Warriors), Greig Laidlaw CAPTAIN (Gloucester)
Stand-off: Finn Russell, Duncan Weir (both Glasgow Warriors)
Centre: Mark Bennett, Peter Horne (Glasgow Warriors), Matt Scott (Edinburgh Rugby), Duncan Taylor (Saracens)
Wing: Sean Lamont, Tommy Seymour (Glasgow Warriors), Tim Visser (Harlequins)
Full-back: Stuart Hogg (Glasgow Warriors), Ruaridh Jackson (Wasps)